Type
Zone
Age range
403.5–401 Ma
Early Devonian
Emsian
Polygnathus inversus
Eocostapolygnathus nothoperbonus
Eocostapolygnathus gronbergi
403.6403.4403.2403.0402.8402.6402.4402.2402.0401.8401.6401.4401.2401.0